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Under oxidative stress conditions, the increasing activities of antioxidant enzymes in plants, such as ascorbate peroxidase (APX), superoxide dismutase (SOD), glutathione reductase (GR), catalase (CAT), peroxidase (POD), peroxiredoxin (PRX) and glutathione S-transferase (GST) participate in the detoxification of peroxides (Mittler, 2002)